Ahrens, D., Monaghan, M. T. & Vogler, A. P. (2007): DNA-based taxonomy for associating adults and larvae in multi-species assemblages of chafers (Coleoptera: Scarabaeidae). Molecular Phylogenetics and Evolution 44(1): 436-449.
Ahrens, D., Zorn, C., GC, Y., Keller, S. & Nagel, P. (2007) Illustrated key of phytophagous scarabs of Nepal. - A Guide to white grubs and chafers of the lower central regions (Coleoptera, Scarabaeidae). Opuscula Biogeographica Basiliensis 5: 1-44.
Sipek, P. & Ahrens, D. (2011) Inferring larval taxonomy and morphology in Maladera species (Coleoptera: Scarabaeidae: Sericini) using DNA taxonomy tools. Systematic Entomology 36: 628–643.